Spermacoce remota

Lam. (1792)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Rubiaceae Genus: Spermacoce

buttonweed

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Description

Key Characters:

Growth Form: Annual or short-lived perennial herbs.

Stems: Stems often purple-tinged, erect or sprawling, 3-10 dm long, glabrous or sparsely retrorsely puberulent in lines on the angles.

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly elliptic, elliptic-lanceolate, or ovate, 1-7 cm long 0.4-3 cm wide. Apex acute to attenuate. Base rounded to cuneate. Blades often purple-tinged, glabrous except puberulent near margins, giving a slightly scabrid texture. Margins with 1 to numerous filiform setae. Petioles 2-4 mm long. Stipules 2-3 mm long, adnate to petioles, forming a sheath, with 5-7 slender, unequal setae 2-5 mm long, stipules of flower-bearing nodes with numerous setae.

Flowers: Flowers numerous in axillary and terminal, dense, sessile clusters, stipule-derived bracteoles with setae 2-3 mm long, subtended by 1-2 or more pairs of bract-like leaves, stipule-derived bracteoles with setae; hypanthium obconical, 2-2.5 mm long. Flowers usually small, bisexual (perfect), sessile, insect-pollinated, often heterostylous. Calyx lobes 2-4(-8) with open aestivation, deltate, ca. 0.6 mm long, ciliate, separated by a fringe. Corolla funnelform or salverform, white or lobes tinged pink, 2.4-2.6 mm long, valvate in bud, the tube usually very slender; nectary disk usually present. Stamens 4, alternate with the corolla lobes, inserted on corolla tube or throat, usually exserted; anthers dithecal, opening by longitudinal slits. Ovary inferior, occasionally partly so, or very rarely superior, 2-celled; ovules 1 per cell, attached to middle of septum; style many as carpels, terminal, slender, filiform, exserted; stigmas lobed or capitate, dry or occasionally wet.

Fruit: Fruit ellipsoid or obovoid-fusiform; 2-4 mm long; pubescent in upper part; dehiscent; the valves bifid; falling completely away. Seeds 2; dull dark brown; oblong-ellipsoid; 1.5-2.5 mm long; transversely sulcate; with well-developed oily endosperm; or endosperm occasionally scanty or absent.

Ploidy:

Habitat: Naturalized and often common in urban areas; along roadsides; and other low elevation disturbed sites.

Elevation Range:

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: Tabl. Encycl. 1: 273 (1792)

Other References

Wagner et al. 1990:1173 (K, O, M, H [as S. assurgens]); Lorence et al. 1995:51/Lorence & Flynn 2006:4/Lorence & Faccenda 2024 (KEY); Oppenheimer 2003:23 (Mo, L); Acevedo-Rodríguez & Strong 2012:847 (Syn. S. assurgens = S. remota)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
2 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
3 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
4 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
5 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
6 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
7 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
8 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
9 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
10 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
11 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
12 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
13 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
14 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
15 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
16 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
17 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
18 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
19 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
20 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
21 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
22 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
23 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
24 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
25 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
26 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
27 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
28 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
29 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
30 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
31 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
32 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
33 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
34 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
35 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
36 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
37 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
38 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
39 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
40 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
41 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
42 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
43 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
44 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
45 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
46 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
47 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
48 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
49 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
50 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
51 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
52 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
53 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
54 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
55 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
56 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
57 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
58 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
59 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
60 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
61 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
62 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
63 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
64 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30
65 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
66 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
67 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
68 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
69 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
70 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
71 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
72 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
73 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
74 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
75 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
76 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
77 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
78 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
79 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
80 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
81 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
82 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
83 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
84 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
85 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
86 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
87 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
88 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
89 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
90 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
91 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
92 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
93 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
94 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
95 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
96 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
97 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
98 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
99 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
100 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
101 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
102 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
103 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
104 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
105 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
106 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
107 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
108 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
109 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
110 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
111 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
112 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
113 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
114 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
115 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
116 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
117 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
118 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
119 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
120 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
121 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
122 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
123 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
124 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
125 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
126 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
127 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
128 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30
129 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
130 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
131 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
132 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
133 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
134 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
135 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
136 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
137 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
138 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
139 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
140 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
141 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
142 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
143 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
144 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
145 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
146 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
147 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
148 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
149 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
150 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
151 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
152 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
153 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
154 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
155 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
156 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
157 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
158 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
159 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
160 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
161 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
162 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
163 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
164 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
165 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
166 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
167 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
168 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
169 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
170 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
171 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
172 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
173 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
174 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
175 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
176 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
177 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
178 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
179 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
180 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
181 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
182 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
183 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
184 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
185 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
186 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
187 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
188 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
189 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
190 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
191 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
192 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30